All 48 in-text bold spans converted: term first-introductions to italics (model merging, genetic drift, continual learning, Fisher-Muller effect, outbreeding depression, directed sex, grounded inheritance/evaluation, and kin), pure-emphasis spans to plain text (the organising-claim sentence, the emergent-null statements, the supported-conclusion sentence). Bold retained only where PNAS uses it: run-in paragraph headings (Design rules. / Analytic tier. / etc.), table labels, and reference volume numbers. Repeat mentions of first-use-italicised terms de-italicised (italics at first use only). 18-pp rebuild clean. Co-Authored-By: Claude Fable 5 <noreply@anthropic.com> Claude-Session: https://claude.ai/code/session_01BkRLcc18rwT2Lysu6PbG7v
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