experiment: E12
kind: speciation
seed: 12
n_replicates: 15

# E12 — MODEL SPECIATION / reproductive isolation (the merge-compatibility limit of the sexual society).
# The Bateson-Dobzhansky-Muller construction: an ancestor; two lineages each substitute a DISJOINT set
# of loci (each parent adaptive, neither carrying an incompatibility); a fraction `rho` of cross-lineage
# locus pairs are incompatibilities (penalty `s`) that only bite when a recombinant inherits BOTH derived
# alleles. Sweeping the divergence d (total substitutions) gives the predicted signature
# COMPATIBLE -> OUTBREEDING DEPRESSION -> HYBRID INVIABILITY, arriving earlier the denser the epistasis
# (rho), with the Orr-Turelli snowball (# incompatibilities ~ (d/2)^2, so fitness falls super-linearly).
# A merged model is a single recombinant (F2-like: hybrid breakdown / recombination load), so this maps
# to postzygotic isolation, not F1 vigour. Falsifier: no outbreeding-depression/isolation progression as
# d and rho grow. Pure seeded NumPy on the E7-E11 genotype machinery (bitwise-reproducible).

speciation:
  landscape: bdm
  L: 20
  rho: [0.1, 0.25, 0.5]       # epistasis DENSITY: fraction of cross-lineage locus pairs that are BDMIs
  divergences: [0, 2, 4, 6, 8, 10, 12, 14, 16, 18, 20]
  s: 1.0                      # incompatibility penalty per realised BDMI
  beta: 1.0                   # additive benefit per derived (adaptive) allele — makes parents fit
  recomb_rate: 0.5            # free recombination (each locus ~ independent parent)
  n_offspring: 500

output:
  dir: results/E12