# LLM-tier model speciation — conflict provokes isolation; duration alone does not E13 carried into language-model weights (0.5B Qwen, LoRA children of one frozen base — the shared base **controls a major source of coordinate mismatch**, allowing a cleaner test of conflict-associated merge failure; averaging can still fail for non-conflict reasons — nonlinear interaction, scaling, capacity — so failures here are *conflict-associated*, not automatically functional). Two knobs, pre-registered readings in the configs; figure `llm_speciation.png` (3 panels; panel B from `results/llm_speciation_add/`). **Design.** Child A: private family `strings`; child B: private family `arith`; shared **ambiguous convention prompts** ("Sort the list […]" — no direction stated) answered *ascending* by A and *descending* by B: each convention harmless alone, contradictory jointly (the Bateson–Dobzhansky– Muller structure). 50/50 soup merge; exact-match verifier; fixed test sets. **Coherence** of a model = max(accuracy under asc grading, under desc grading) on the shared prompts — a coherent parent scores under its own convention; a hybrid mixing conventions scores low under both (the `μ(S)/2` floor made operational). ### Finding 1 — function-specific hybrid breakdown (the conflict knob; panel A) Once the conventions are trained (`conflict_frac ≥ 0.25`), each parent performs under its own convention (~0.20–0.25 — low absolute values: sorting long lists is hard for 0.5B) while the merge's coherence sits at **0.02–0.08, below BOTH parents under either grading** — the hybrid loses precisely the conflicted function. (At `frac = 0` no one has seen the ambiguous prompts; that point is a no-conflict baseline, not part of the cliff.) ### Finding 2 — the damage does not spread (the de-confounded `add` design; panel B) In the original ("replace") sweep, higher conflict fraction mechanically means *less private-family training*, so the merge's private-family decline is confounded. The **`add` design** (`llm_speciation_add`, 3 seeds) holds each child's private training fixed and adds conflict data on top: the merge's private-family accuracy then shows **no trend with conflict** (0.74–0.88, tracking parent A's 0.82–0.87 within seed noise at every level). Conflict damage is **localised to the conflicted function**; it does not corrupt the disjoint skills — at this scale, hybrid breakdown is surgical, not global. Honest rider: 0.5B soup merges carry large *intrinsic* seed variance even at zero conflict (sd up to 0.28) — the same averaging-fragility seen in `llm_moe_hard_seeds`. ### Finding 3 — the duration null: over-specialisation does not erode mergeability (panel C) Pure disjoint specialists over-trained from 1 to 12 epochs (no shared data at all): the merged model *improves* (0.84 → 0.94 mean-private) and stays **above the best parent at every duration**. The MLP tier's "no emergent isolation" null (`speciation_real_emergent`) **generalises to LLM weights** in this regime — relevant to the report that averaging prefers under-trained experts (arXiv:2607.11997): in our disjoint-family setting, no such over-training penalty appears; the theory's prediction is that their effect should trace to *conflicting conventions on shared circuitry*, which the `epistasis_predicts` experiment (work order) will test directly. **Speciation across all three tiers now reads:** analytic (E12: cliff, epistasis-dependence, snowball) → MLP (E13: functional residual survives the full symmetry group; no emergent isolation) → LLM (this run: function-specific hybrid breakdown under conflict; no isolation from duration or specialisation alone). Isolation must be provoked by functional conflict at every tier tested. Falsifiers (not triggered): merge coherence matching the parents (no breakdown), or merged private-family accuracy declining with conflict in the `add` design (global corruption).