# kernel — the learning kernel: why real learners deviate from neutral drift *(This legend covers both `results/figS2_kernel_sharpen/` and `results/figS2_kernel_smooth/`; the figure `kernel.png` is written into both.)* **Claim tested.** Neutral Wright–Fisher drift (the histogram bridge, and the baseline of Riis 2026) is the *null* model of collapse. But Layer 1.5 showed real trained models deviate from it — and in **opposite directions**. Can a single extension of Layer 1 — a parameterized *learning kernel* on the refit step, `p_{t+1} = T_θ(counts/n)` — reproduce both deviations, and does neutral drift genuinely fail without it? **Setup.** The kernel (`knowledge/kernel.py`) has two population-genetics knobs, both reducing to neutral drift at their defaults (so the histogram and every scientific-validation test are unchanged): **reset `u`** — mutation toward a prior (`p ← (1−u)p + u·π`), i.e. smoothing; and **temperature `τ`** — sharpening (`p ∝ p^{1/τ}`, `τ<1` concentrates), i.e. mode-competition. Two matched-to-neural regimes, 24 replicates each. ### The four panels (`kernel.png`; blue = neutral, red = kernel-on, green dashed = the real neural model) **Top row — VAE regime (`n=6000`, `K=30`), pro-collapse:** 1. **Heterozygosity.** Neutral drift is **inert** — at `n=6000` it barely moves (`H` stays at `H*`). Yet the real VAE (green) collapsed to `H≈0`. **Sharpening (`τ=0.8`) reproduces the collapse.** Neutral drift is *falsified*; the estimator's mode-competition is required. 2. **Support.** Neutral holds ~all 30 modes; sharpening → 1 mode, matching the VAE. **Bottom row — RNN regime (`n=200`, `K=256`), anti-collapse:** 3. **Heterozygosity.** Neutral drift drives `H → 0`, but the real RNN (green) only *partially* collapses (`H` floors at ~0.68). **Mutation `u=0.006` reproduces the floor.** The estimator here *removes* collapse pressure. 4. **Forward-KL.** Neutral diverges; smoothing plateaus. Honest caveat: uniform-mutation plateaus *above* the RNN's KL (~5 vs ~2) — evidence the RNN's smoothing target is **truth-like, not uniform** (a refinement for future work). The *sign* is unambiguous. ### Takeaway Model collapse in real learners = **neutral drift ⊕ an architecture-specific estimator-bias operator that can point either way.** The histogram sits at the neutral null (`u=0, τ=1`); the VAE **sharpens** (adds collapse); the RNN/MLP **smooth** (add a diversity floor). This mechanistically explains the Layer-1.5 architecture-generality result and the softened neural `g*`, and develops the exact axis Riis (2026) names as future work ("different smoothing schemes… each induce their own fixed-point geometry… a natural direction for further work"). `u`/`τ` are calibrated from a single neural diagnostic and pinned in the configs. **Falsifier (not triggered):** if neutral drift had already reproduced the neural curves, the estimator axis would be superfluous — instead it fails in *both* regimes, oppositely.